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Divides observed mutations by the number of sites screened and generations to give the per-site per-generation mutation rate.
Mutation rate
Rate = mutations observed / (sites screened x generations); mutations per genome = rate x genome size
About 1.2 x 10^-8 per site per generation, giving roughly 50 to 70 new mutations per genome.
Sperm continue dividing throughout life, so the paternal mutation contribution rises with age.